Prehistoric humans venerated skulls and it is difficult for modern humans in their context to understand why they did so. Here is why.
When humans first became self aware, separating themselves in level of consciousness from animals, one of the first things they learned about themselves was that they recognized the mechanics of eyesight. By mechanics I mean that humans thought about how when they shut their eyes they cannot see and when they open their eyes they can see. Animals do not think about that, they just do it. Humans realized that all of life required sight and so it was to them the center of their being. So when animals and humans died they were curious as to what was in the skulls and also venerated it as the center of their ability to survive. (Remember that prehistoric man required eye sight to forage for food).
It is only much later in history that skulls became gory and dark, or trophies with death connotations. Early humans viewed skulls as the place that housed their vision. So they were kind of like lighthouse symbols to them, rather than symbols of death. They recognized that animals also have eyes in their head, just like them, and oxen had to see in order to walk and help humans plow. So skulls were of great interest and veneration to prehistoric humans because they were acknowledging the mystery of sight and the feeling of "existing behind one's eyeballs" as they became self aware and starting thinking about these things and wondering.
Yeah, I know lots of interesting things... just don't brag about it like some I could mention.